Harpalus (beetle)

Harpalus
Harpalus affinis
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Family:	Carabidae
Subfamily:	Harpalinae
Tribe:	Harpalini
Genus:	Harpalus Latreille, 1802
Synonyms
Harpalellus Lindroth, 1968 Harpalobrachys Pseudoophonus Motschoulsky, 1844

Harpalus is a genus of ground beetle first described by Pierre André Latreille in 1802.

Species

Harpalus contains the following 430 species:

Subgenus Cephalophonus Ganglbauer, 1891

Harpalus cephalotes Fairmaire & Laboulbène, 1854

Subgenus Cryptophonus Brandmayr & Zetto Brandmayr, 1982

Harpalus agnatus Reiche, 1850

Harpalus cyrenaicus Koch, 1939

Harpalus fulvus Dejean, 1829

Harpalus grilli Kataev, 2002

Harpalus idiotus Bates, 1889

Harpalus janinae Jeanne, 1984

Harpalus litigiosus Dejean, 1829

Harpalus melancholicus Dejean, 1829

Harpalus schaumii Wollaston, 1864

Harpalus tenebrosus Dejean, 1829

Subgenus Glanodes Casey, 1914

Harpalus cohni Ball, 1972

Harpalus corpulentus (Casey, 1914)

Harpalus huachuca Ball, 1972

Harpalus obliquus G.Horn, 1880

Harpalus puncticeps (Casey, 1914)

Harpalus stephani Ball, 1972

Subgenus Harpalus Latreille, 1802

Harpalus acupalpoides Reitter, 1900

Harpalus aeneipennis (Faldermann, 1836)

Harpalus aequicollis Motschulsky, 1844

Harpalus affinis (Schrank, 1781)

Harpalus agakhaniantzi (Mikhailov, 1972)

Harpalus akbensis Jedlicka, 1958

Harpalus akinini Tschitscherine, 1895

Harpalus alajensis Tschitscherine, 1898

Harpalus albanicus Reitter, 1900

Harpalus alexandrae Kataev, 1990

Harpalus alexeevi Kataev, 1990

Harpalus alpivagus Tschitscherine, 1899

Harpalus amarellus Bates, 1891

Harpalus amariformis Motschulsky, 1844

Harpalus amplicollis Ménétriés, 1848

Harpalus amputatus Say, 1830

Harpalus anatolicus Tschitscherine, 1898

Harpalus angulatus Putzeys, 1878

Harpalus angustipennis Boheman, 1848

Harpalus angustitarsis Reitter, 1887

Harpalus animosus Casey, 1924

Harpalus anisodactyliformis Solsky, 1874

Harpalus antonowi (Tschitscherine, 1898)

Harpalus anxioides Kataev, 1991

Harpalus anxius (Duftschmid, 1812)

Harpalus araraticus Mlynar, 1979

Harpalus arcuatus Tschitscherine, 1898

Harpalus arnoldii Kataev, 1988

Harpalus ascetes Kataev & Wrase, 1997

Harpalus asemus Basilewsky, 1946

Harpalus aterrimus Casey, 1914

Harpalus atratus Latreille, 1804

Harpalus atrichatus Hatch, 1949

Harpalus atripes Casey, 1914

Harpalus attenuatus Stephens, 1828

Harpalus australasiae Dejean, 1829

Harpalus autumnalis (Duftschmid, 1812)

Harpalus badakschanus Jedlicka, 1956

Harpalus baleensis Clarke, 1973

Harpalus balli Noonan, 1991

Harpalus basanicus J.Sahlberg, 1913

Harpalus basilewskyi Facchini, 2003

Harpalus basuto Basilewsky, 1958

Harpalus bellieri Reiche, 1861

Harpalus beneshi Kataev & Wrase, 1997

Harpalus brachypterus Tschitscherine, 1898

Harpalus brevicornis Germar, 1823

Harpalus brevis Motschulsky, 1844

Harpalus breviusculus Chaudoir, 1846

Harpalus bucharicus Tschitscherine, 1898

Harpalus bulirschi (Facchini, 2012)

Harpalus bungii Chaudoir, 1844

Harpalus caeruleatus Bates, 1878

Harpalus caiphus Reiche & Saulcy, 1855

Harpalus calathoides Motschulsky, 1844

Harpalus capicola Dejean, 1829

Harpalus cardoni Antoine, 1922

Harpalus caspius (Steven, 1806)

Harpalus cautus Dejean, 1829

Harpalus chalcentus Bates, 1873

Harpalus chasanensis Lafer, 1989

Harpalus chobautianus Lutshnik, 1922

Harpalus chrysopus Reitter, 1887

Harpalus circumpunctatus Chaudoir, 1846

Harpalus cisteloides Motschulsky, 1844

Harpalus clarkei Kataev & J.Schmidt, 2020

Harpalus compressus Motschulsky, 1844

Harpalus contemptus Dejean, 1829

Harpalus convexus Faldermann, 1836

Harpalus corporosus (Motschulsky, 1862)

Harpalus corrugatus Basilewsky, 1958

Harpalus crates Bates, 1883

Harpalus cupreus Dejean, 1829

Harpalus cursorius Péringuey, 1896

Harpalus cyanopterus (Tschitscherine, 1897)

Harpalus cyclogonus Chaudoir, 1844

Harpalus danieli Reitter, 1900

Harpalus davidianus Tschitscherine in Jakobson, 1903

Harpalus decipiens Dejean, 1829

Harpalus defector Péringuey, 1896

Harpalus desertus LeConte, 1859

Harpalus diligens Tschitscherine, 1898

Harpalus dimidiatus (P.Rossi, 1790)

Harpalus dispar Dejean, 1829

Harpalus dissitus Antoine, 1931

Harpalus distinguendus (Duftschmid, 1812)

Harpalus diversicollis Basilewsky, 1958

Harpalus dubius Boheman, 1848

Harpalus dudkoi Kataev, 2011

Harpalus durangoensis Bates, 1891

Harpalus ebeninus Heyden, 1870

Harpalus egenus Dejean, 1829

Harpalus egorovi Lafer, 1989

Harpalus ellipsis LeConte, 1847

Harpalus elliptipennis Facchini, 2003

Harpalus erosus Mannerheim, 1825

Harpalus exiguus Boheman, 1848

Harpalus famelicus Tschitscherine, 1898

Harpalus farkaci Kataev & Wrase, 1995

Harpalus fimetarius Dejean, 1829

Harpalus flavescens (Piller & Mitterpacher, 1783)

Harpalus flavicornis Dejean, 1829

Harpalus foveiger Tschitscherine, 1895

Harpalus franzi Mateu, 1954

Harpalus frater Chaudoir, 1876

Harpalus fraternus LeConte, 1852

Harpalus froelichii Sturm, 1818

Harpalus fulvicornis Thunberg, 1806

Harpalus fulvilabris Mannerheim, 1853

Harpalus fulvipennis Chaudoir, 1843

Harpalus fuscicornis Ménétriés, 1832

Harpalus fuscipalpis Sturm, 1818

Harpalus fuscipennis Wiedemann, 1825

Harpalus fuscoaeneus Dejean, 1829

Harpalus ganssuensis Semenov, 1889

Harpalus giacomazzoi Kataev & Wrase, 1996

Harpalus gilgil Basilewsky, 1946

Harpalus gisellae Csiki, 1932

Harpalus glasunovi Kataev, 1987

Harpalus gravis LeConte, 1858

Harpalus gregoryi Alluaud, 1917

Harpalus hartmanni Kataev, 2002

Harpalus herbivagus Say, 1823

Harpalus heyrovskyi Jedlicka, 1928

Harpalus hiekei Kataev & Wrase, 2010

Harpalus hirtipes (Panzer, 1796)

Harpalus honestus (Duftschmid, 1812)

Harpalus hospes Sturm, 1818

Harpalus hybridus Boheman, 1848

Harpalus impressus Roth, 1851

Harpalus inconcinnus Chaudoir, 1876

Harpalus indianus Csiki, 1932

Harpalus indicola Bates, 1878

Harpalus indigens Casey, 1924

Harpalus inexspectatus Kataev, 1989

Harpalus ingenuus Tschitscherine, 1898

Harpalus innocuus LeConte, 1863

Harpalus italus Schaum, 1860

Harpalus jeanneli Basilewsky, 1946

Harpalus jordanus Jedlicka, 1964

Harpalus kabakianus Kataev, 1988

Harpalus kabakovi Kataev, 1987

Harpalus kadleci Kataev & Wrase, 1995

Harpalus kadyrbekovi Kataev, 1988

Harpalus kagyzmanicus Kataev, 1984

Harpalus kandaharensis Jedlicka, 1956

Harpalus karakorum Jedlicka, 1958

Harpalus karamani Apfelbeck, 1902

Harpalus kazanensis Jedlicka, 1958

Harpalus kaznakovi Kataev & Wrase, 1997

Harpalus kibonoti Alluaud, 1927

Harpalus kirgisicus (Motschulsky, 1844)

Harpalus kiritshenkoi Kataev, 1990

Harpalus klapperichi Jedlicka, 1956

Harpalus kmecoi Facchini, 2003

Harpalus kozlovi Kataev, 1993

Harpalus kryzhanovskii Kataev, 1988

Harpalus kunarensis Kataev, 1993

Harpalus laetus Reiche, 1843

Harpalus laevipes Zetterstedt, 1828

Harpalus lama Kataev & Wrase, 1997

Harpalus lateralis Dejean, 1829

Harpalus laticeps LeConte, 1850

Harpalus latus (Linnaeus, 1758)

Harpalus lederi Tschitscherine, 1899

Harpalus lethierryi Reiche, 1860

Harpalus lewisii LeConte, 1865

Harpalus longipalmatus Mordkovitsh, 1969

Harpalus lopezi J.Serrano & Lencina, 2009

Harpalus lugubris Boheman, 1848

Harpalus lumbaris Mannerheim, 1825

Harpalus luteicornis (Duftschmid, 1812)

Harpalus lutshniki Schauberger, 1932

Harpalus macronotus Tschitscherine, 1893

Harpalus mairei Peyerimhoff, 1928

Harpalus major (Motschulsky, 1850)

Harpalus makhekensis Basilewsky, 1958

Harpalus manas Kataev, 1990

Harpalus marginellus Gyllenhal, 1827

Harpalus martini Van Dyke, 1926

Harpalus masoreoides Bates, 1878

Harpalus massarti Burgeon, 1936

Harpalus mauritanicus Gaubil, 1844

Harpalus medvedevi Kataev, 2006

Harpalus megacephalus LeConte, 1847

Harpalus melaneus Bates, 1878

Harpalus metallinus Ménétriés, 1836

Harpalus metarsius Andrewes, 1930

Harpalus meteorus Basilewsky, 1946

Harpalus michaili Kataev, 1990

Harpalus michailovi Kataev, 1987

Harpalus microthorax (Motschulsky, 1849)

Harpalus miles Péringuey, 1896

Harpalus minutissimus Facchini, 2015

Harpalus minutulus Kataev & Liang, 2004

Harpalus mitridati Pliginskiy, 1915

Harpalus mlynari Kataev, 1990

Harpalus modestus Dejean, 1829

Harpalus morvani Kataev, 2002

Harpalus murzini Kataev, 2008

Harpalus nanniscus Péringuey, 1896

Harpalus natalensis Boheman, 1848

Harpalus natalicus Péringuey, 1896

Harpalus neglectus Audinet-Serville, 1821

Harpalus nevadensis K. & J.Daniel, 1898

Harpalus nigrans A.Morawitz, 1862

Harpalus nigritarsis C.R.Sahlberg, 1827

Harpalus numidicus Bedel, 1893

Harpalus nyassicus Basilewsky, 1946

Harpalus oblitus Dejean, 1829

Harpalus obnixus Casey, 1924

Harpalus ochropus Kirby, 1837

Harpalus oodioides Dejean, 1829

Harpalus opacipennis (Haldeman, 1843)

Harpalus optabilis Dejean, 1829

Harpalus ovtshinnikovi Kataev, 1990

Harpalus pallidipennis A.Morawitz, 1862

Harpalus pallipes Chaudoir, 1837

Harpalus parallelocollis Facchini, 2003

Harpalus parasinuatus Kataev & Liang, 2007

Harpalus parvulus Dejean, 1829

Harpalus petri Tschitscherine, 1902

Harpalus picipennis (Duftschmid, 1812)

Harpalus plancyi Tschitscherine, 1897

Harpalus plenalis Casey, 1914

Harpalus politus Dejean, 1829

Harpalus polyglyptus Schaum, 1862

Harpalus potanini Tschitscherine, 1906

Harpalus praecurrens Schauberger, 1934

Harpalus procognatus Lorenz, 1998

Harpalus progrediens Schauberger, 1922

Harpalus prosperus Basilewsky, 1972

Harpalus providens Casey, 1914

Harpalus pseudoserripes Reitter, 1900

Harpalus pterostichus (Reitter, 1900)

Harpalus puetzi Kataev & Wrase, 1997

Harpalus pulchrinulus Reitter, 1900

Harpalus pulvinatus Ménétriés, 1848

Harpalus pumilus Sturm, 1818

Harpalus punctatostriatus Dejean, 1829

Harpalus punctipennis Mulsant, 1852

Harpalus pusillus (Motschulsky, 1850)

Harpalus pygmaeus Dejean, 1829

Harpalus quadratus Chaudoir, 1846

Harpalus raphaili Kataev, 1997

Harpalus reflexus Putzeys in Schneider & Leder, 1878

Harpalus reitteri Wrase & Kataev, 2011

Harpalus remboides Solsky, 1874

Harpalus reversus Casey, 1924

Harpalus rhemboides Solsky, 1874

Harpalus rivalsi Jeannel, 1948

Harpalus rotundus Facchini, 2003

Harpalus rougemonti Clarke, 1973

Harpalus rubripes (Duftschmid, 1812)

Harpalus rufipalpis Sturm, 1818

Harpalus rufiscapus Gebler, 1833

Harpalus rufitarsoides Schauberger, 1934

Harpalus rufomarginatus (Boheman, 1848)

Harpalus rumelicus Apfelbeck, 1904

Harpalus salinulus Reitter, 1900

Harpalus salinus Dejean, 1829

Harpalus sanctaehelenae Basilewsky, 1972

Harpalus saxicola Dejean, 1829

Harpalus semenowi Tschitscherine, 1901

Harpalus semipunctatus Dejean, 1829

Harpalus serripes (Quensel, 1806)

Harpalus servus (Duftschmid, 1812)

Harpalus setiporus (Reitter, 1894)

Harpalus siculus Dejean, 1829

Harpalus silipes Dejean, 1831

Harpalus sinuaticollis Facchini, 2015

Harpalus sinuatus Tschitscherine, 1893

Harpalus smaragdinus (Duftschmid, 1812)

Harpalus smyrnensis Heyden, 1888

Harpalus solitaris Dejean, 1829

Harpalus somereni Basilewsky, 1946

Harpalus somnulentus Dejean, 1829

Harpalus spadiceus Dejean, 1829

Harpalus spretus Péringuey, 1896

Harpalus spurius Péringuey, 1896

Harpalus stevenii Dejean, 1829

Harpalus stevensi Kataev, 2011

Harpalus stoetznerianus Schauberger, 1932

Harpalus strenuus Tschitscherine, 1898

Harpalus subaeneus Boheman, 1848

Harpalus subcylindricus Dejean, 1829

Harpalus subtruncatus Chaudoir, 1846

Harpalus sulphuripes Germar, 1823

Harpalus sundaicus Schauberger, 1933

Harpalus sushenicus Kataev, 1990

Harpalus suturangulus Reitter, 1887

Harpalus szalliesi Kataev & Wrase, 1995

Harpalus taciturnus Dejean, 1829

Harpalus tangutorum Kataev, 1993

Harpalus tardus (Panzer, 1796)

Harpalus tarsalis Mannerheim, 1825

Harpalus terrestris (Motschulsky, 1844)

Harpalus tibeticus Andrewes, 1930

Harpalus tichonis Jakobson, 1907

Harpalus tiridates Reitter, 1900

Harpalus tithonus Reitter, 1900

Harpalus tjanschanicus Semenov, 1889

Harpalus torosensis Jedlicka, 1961

Harpalus torridoides Reitter, 1900

Harpalus trichophorus Tschitscherine, 1897

Harpalus triseriatus A.Fleischer, 1897

Harpalus turcicus Jedlicka, 1958

Harpalus turmalinus Erichson, 1847

Harpalus udege Lafer, 1989

Harpalus uniformis (Motschulsky, 1844)

Harpalus ussuricus Mlynar, 1979

Harpalus vanemdeni Schauberger, 1932

Harpalus venator Boheman, 1848

Harpalus ventralis LeConte, 1847

Harpalus vereschaginae Kataev, 1988

Harpalus vernicosus Kataev & Liang, 2007

Harpalus viridanus (Motschulsky, 1844)

Harpalus viridicupreus Reiche, 1843

Harpalus vittatus Gebler, 1833

Harpalus wadiensis Jedlicka, 1964

Harpalus wohlberedti Emden, 1932

Harpalus xanthopus Gemminger & Harold, 1868

Harpalus zabroides Dejean, 1829

Harpalus zhdankoi Kataev, 1990

Subgenus Loboharpalus Schauberger, 1932

Harpalus platynotus Bates, 1873

Harpalus rubefactus Bates, 1873

Subgenus Opadius Casey, 1914

Harpalus apache Kataev, 2010

Harpalus cordatus (LeConte, 1853)

Harpalus cordifer Notman, 1919

Subgenus Plectralidus Casey, 1914

Harpalus erraticus Say, 1823

Harpalus retractus LeConte, 1863

Subgenus Pseudoophonus Motschulsky, 1844

Harpalus actiosus Casey, 1914

Harpalus aenigma (Tschitscherine, 1897)

Harpalus alienus Bates, 1878

Harpalus aogashimensis (Habu, 1957)

Harpalus azumai Habu, 1968

Harpalus babai Habu, 1973

Harpalus calceatus (Duftschmid, 1812)

Harpalus caliginosus (Fabricius, 1775)

Harpalus capito A.Morawitz, 1862

Harpalus compar LeConte, 1847

Harpalus coreanus (Tschitscherine, 1895)

Harpalus davidi (Tschitscherine, 1897)

Harpalus disimuciulus Huang; Lei; Yan & Hu, 1996

Harpalus eous Tschitscherine, 1901

Harpalus erythropus Dejean, 1829

Harpalus faunus Say, 1823

Harpalus fokienensis Schauberger, 1930

Harpalus griseus (Panzer, 1796)

Harpalus hatchi Ball & Anderson, 1962

Harpalus hauserianus Schauberger, 1929

Harpalus indicus Bates, 1891

Harpalus jureceki (Jedlicka, 1928)

Harpalus katiae F.Battoni, 1985

Harpalus liobasis Chaudoir, 1868

Harpalus meghalayensis Kataev, 2001

Harpalus meridianus Andrewes, 1923

Harpalus paratus Casey, 1924

Harpalus pastor Motschulsky, 1844

Harpalus pensylvanicus (DeGeer, 1774)

Harpalus poncei Will, 2002

Harpalus protractus Casey, 1914

Harpalus pseudohauserianus Kataev, 2001

Harpalus pseudophonoides Schauberger, 1930

Harpalus roninus Bates, 1873

Harpalus rufipes (DeGeer, 1774)

Harpalus sericatus Tschitscherine, 1906

Harpalus simplicidens Schauberger, 1929

Harpalus singularis Tschitscherine, 1906

Harpalus sinicus Hope, 1845

Harpalus suensoni Kataev, 1997

Harpalus texanus Casey, 1914

Harpalus tridens A.Morawitz, 1862

Harpalus ussuriensis Chaudoir, 1863

Harpalus vagans LeConte, 1865

Harpalus yinchuanensis Huang, 1993

Subgenus Semiophonus Schauberger, 1933

Harpalus signaticornis (Duftschmid, 1812)

Subgenus Zangoharpalus Huang, 1998

Harpalus mariae Kataev, 1997

Harpalus microdemas Schauberger, 1932

Harpalus praticola Bates, 1891

Harpalus pseudotinctulus Schauberger, 1932

Harpalus tinctulus Bates, 1873

Extinct, not assigned to subgenus

†Harpalus abolitus C. & L.Heyden, 1866

†Harpalus anactus Giebel, 1856

†Harpalus bruckmanni Heer, 1862

†Harpalus burmeisteri Giebel, 1856

†Harpalus conditus Scudder, 1900

†Harpalus constrictus Heer, 1862

†Harpalus deletus Oustalet, 1864

†Harpalus dilatatus Theobald, 1937

†Harpalus ewaldi Giebel, 1856

†Harpalus excavatus Forster, 1891

†Harpalus heeri Giebel, 1856

†Harpalus knorri Giebel, 1856

†Harpalus lheritieri Piton, 1940

†Harpalus liasinus Giebel, 1856

†Harpalus maceratus Wickham, 1911

†Harpalus nero Oustalet, 1864

†Harpalus nitens Birket-Smith, 1977

†Harpalus nuperus Scudder, 1900

†Harpalus obtusus Forster, 1891

†Harpalus oustaleti Meunier, 1915

†Harpalus pleistocenicus Lomnicki, 1894

†Harpalus redivivus Wickham, 1917

†Harpalus schlotheimi Giebel, 1856

†Harpalus sinis Heer, 1847

†Harpalus stierlini Heer, 1862

†Harpalus stygius Heer, 1862

†Harpalus tabidus Heer, 1847

†Harpalus tardigradus Heer, 1862

†Harpalus ulomaeformis Wickham, 1917

†Harpalus veterum Cockerell, 1920

†Harpalus whitfieldi Scudder, 1900